Footnotes

• Handling editor Anna Maria Geretti

• Contributors DR designed this study; AR and ST collected the data; AR analysed the data; and AR, ST, KN, MT, MD, JW and DR all contributed to the final manuscript.

• Funding The authors have not declared a specific grant for this research from any funding agency in the public, commercial or not-for-profit sectors.

• Competing interests None declared.

• Provenance and peer review Not commissioned; internally peer reviewed.